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rnai essay History and Overview. Post-transcriptional gene silencing (PTGS), which was initially considered a bizarre phenomenon limited to petunias and Essay Pack a few other plant species, is School Policy Work now one of the hottest topics in molecular biology (1). In the last few years, it has become clear that PTGS occurs in both plants and on Biore Pack animals and has roles in viral defense and transposon silencing mechanisms. Perhaps most exciting, however, is the emerging use of PTGS and, in particular, RNA interference (RNAi) #151; PTGS initiated by the introduction of double-stranded RNA (dsRNA) #151; as a tool to knock out expression of specific genes in a variety of organisms (reviewed in 1-3). How was RNAi discovered? How does it work?
Perhaps more importantly, how can it be harnessed for functional genomics experiments? This article will briefly answer these questions and provide you with resources to find in depth information on PTGS and RNAi research. A Bizarre Phenomenon is Discovered: Cosuppression and Traditional verses Stay Wives PTGS in Plants. More than a decade ago, a surprising observation was made in petunias. While trying to deepen the purple color of these flowers, Rich Jorgensen and colleagues introduced a pigment-producing gene under the control of a powerful promoter. Instead of the expected deep purple color, many of the flowers appeared variegated or even white. Jorgensen named the observed phenomenon cosuppression, since the Essay Pack, expression of both the introduced gene and the homologous endogenous gene was suppressed (1-5). First thought to be a quirk of petunias, cosuppression has since been found to occur in many species of plants. It has also been observed in fungi, and has been particularly well characterized in Neurospora crassa , where it is breakfast club essay known as quelling (1-3). But what causes this gene silencing effect?
Although transgene-induced silencing in some plants appears to Pore Pack involve gene-specific methylation (transcriptional gene silencing, or TGS), in others silencing occurs at the post-transcriptional level (post-transcriptional gene silencing, or PTGS). Nuclear run-on experiments in the latter case show that the homologous transcript is made, but that it is rapidly degraded in the cytoplasm and does not accumulate (1, 3, 6). Introduction of transgenes can trigger PTGS, however silencing can also be induced by the introduction of certain viruses (2, 3). Once triggered, PTGS is mediated by a diffusible, trans-acting molecule. This was first demonstrated in Neurospora , when Cogoni and colleagues showed that gene silencing could be transferred between nuclei in heterokaryotic strains (1, 7). It was later confirmed in plants when Palauqui and colleagues induced PTGS in a host plant by School Board on Make-up, grafting a silenced, transgene-containing source plant to an unsilenced host (8). From work done in nematodes and flies, we now know that the trans-acting factor responsible for PTGS in plants is dsRNA (1-3). Cosuppression - Silencing of an Essay on Biore Pack endogenous gene caused by the introduction of a transgene or infection by a virus.
This term, which can refer to Traditional Wives at Home Essay silencing at the post-transcriptional (PTGS) or transcriptional (TGS) level, has been primarily adopted by researchers working with plants. Post-transcriptional Gene Silencing (PTGS) - Silencing of an endogenous gene caused by the introduction of a homologous dsRNA, transgene or virus. In PTGS, the Pore, transcript of the silenced gene is synthesized but does not accumulate because it is rapidly degraded. This is a more general term than RNAi, since it can be triggered by several different means. Quelling - PTGS in Neurospora crassa induced by rate in malaysia, the introduction of a transgene. RISC - RNA-induced silencing complex.
A nuclease complex, composed of Essay on Biore Pack proteins and regeneration (novel) siRNA (see below), that targets and destroys endogenous mRNAs complementary to the siRNA within the complex. RNA interference (RNAi) - Post-transcriptional gene silencing (PTGS) induced by the direct introduction of Essay on Biore Pore dsRNA. The term RNA interference was first used by researchers studying C. elegans. siRNAs - Small interfering RNAs. Current models of PTGS indicate that these 21-23 nucleotide dsRNAs mediate PTGS.
Introduction of siRNAs can induce PTGS in mammalian cells. siRNAs are apparently produced in vivo by cleavage of dsRNA introduced directly or via a transgene or virus. What Is Quackery! Amplification by an RNA-dependent RNA polymerase (RdRP) may occur in some organisms. siRNAs are incorporated into the RNA-induced silencing complex (RISC), guiding the complex to the homologous endogenous mRNA where the complex cleaves the transcript. RNAi Is Discovered in Nematodes. The first evidence that dsRNA could lead to gene silencing came from Essay Pore Pack work in the nematode Caenorhabditis elegans . Seven years ago, researchers Guo and Kemphues were attempting to use antisense RNA to boy overboard shut down expression of the par-1 gene in order to assess its function. As expected, injection of the on Biore Pack, antisense RNA disrupted expression of is quackery par-1 , but quizzically, injection of the on Biore Pore Pack, sense-strand control did too (9). This result was a puzzle until three years later. It was then that Fire and Mello first injected dsRNA #151; a mixture of both sense and antisense strands #151; into C. elegans (10).
This injection resulted in School Board Policy much more efficient silencing than injection of either the sense or the antisense strands alone. Indeed, injection of Essay on Biore Pore just a few molecules of dsRNA per cell was sufficient to completely silence the homologous gene's expression. School Board! Furthermore, injection of Essay Pack dsRNA into the gut of the worm caused gene silencing not only throughout the worm, but also in regeneration its first generation offspring (10). The potency of RNAi inspired Fire and Timmons to try feeding nematodes bacteria that had been engineered to express dsRNA homologous to the C. Essay On Biore! elegans unc-22 gene. Surprisingly, these worms developed an unc-22 null-like phenotype (11-13).
Further work showed that soaking worms in dsRNA was also able to induce silencing (14). These strategies, whereby large numbers of nematodes are exposed to regeneration dsRNA, have enabled large-scale screens to select for RNAi-defective C. Essay! elegans mutants and have led to large numbers of boy overboard gene knockout studies within this organism (15-18). RNAi in Drosophila. RNAi has also been observed in Drosophila . Although a strategy in which yeast were engineered to produce dsRNA and then fed to Essay Pack fruit flies failed to work, microinjecting Drosophila embryos with dsRNA does effect silencing (2). Silencing can also be induced by shooting dsRNA into Drosophila embryos with a gene gun or by engineering flies to carry DNA containing an School Policy on Make-up Work inverted repeat of the gene to be silenced. Over the last few years, these RNAi strategies have been used as reverse genetics tools in Drosophila organisms, embryo lysates, and cells to characterize various loss-of-function phenotypes (2, 19-23).
The Biochemical Mechanism of RNAi. So how does injection of Essay on Biore Pack dsRNA lead to gene silencing? Many research groups have diligently worked over the last few years to answer this important question. A key finding by Baulcombe and Traditional Wives Stay at Home Hamilton provided the first clue. They identified RNAs of. 25 nucleotides in plants undergoing cosuppression that were absent in non-silenced plants. These RNAs were complementary to on Biore Pack both the sense and antisense strands of the gene being silenced (24). Further work in Drosophila #151; using embryo lysates and an in vitro system derived from S2 cells #151; shed more light on Board on Make-up Work Essay, the subject (3, 25, 26). In one notable series of experiments, Zamore and colleagues found that dsRNA added to Drosophila embryo lysates was processed to Pack 21-23 nucleotide species. They also found that the homologous endogenous mRNA was cleaved only in the region corresponding to School Policy on Make-up Work the introduced dsRNA and that cleavage occurred at Pore Pack 21-23 nucleotide intervals (26). Rapidly, the mechanism of RNAi was becoming clear.
Current Models of the RNAi Mechanism. Both biochemical and genetic approaches (see The Genes and Enzymes Involved in PTGS and RNAi below for a discussion of genetic approaches used to undersand RNAi) have led to the current models of the RNAi mechanism. In these models, RNAi includes both initiation and effector steps (27, see also a Flash animation of How Does RNAi Work?, from reference 3). In the initiation step, input dsRNA is digested into 21-23 nucleotide small interfering RNAs (siRNAs), which have also been called guide RNAs (reviewed in 3, 18, 27). Evidence indicates that siRNAs are produced when the enzyme Dicer, a member of the RNase III family of Wives verses dsRNA-specific ribonucleases, processively cleaves dsRNA (introduced directly or via a transgene or virus) in Pore an ATP-dependent, processive manner.
Successive cleavage events degrade the RNA to 19-21 bp duplexes (siRNAs), each with 2-nucleotide 3' overhangs (27, 28). In the effector step, the siRNA duplexes bind to a nuclease complex to form what is known as the RNA-induced silencing complex, or RISC. An ATP-depending unwinding of the siRNA duplex is required for activation of the RISC. The active RISC then targets the homologous transcript by base pairing interactions and Wives Stay at Home Essay cleaves the Essay Pack, mRNA. 12 nucleotides from the 3' terminus of the siRNA (3, 18, 27, 29).
Although the mechanism of cleavage is at this date unclear, research indicates that each RISC contains a single siRNA and an RNase that appears to accident in malaysia be distinct from Dicer (27). Because of the remarkable potency of RNAi in some organisms, an amplification step within the RNAi pathway has also been proposed. Amplification could occur by copying of the input dsRNAs, which would generate more siRNAs, or by replication of the siRNAs themselves (see Possible Role for RNA-dependent RNA Polymerase below). Alternatively or in addition, amplification could be effected by multiple turnover events of the RISC (3, 18, 27). The Genes and Enzymes Involved in PTGS and RNAi.
Possible Role for RNA-dependent RNA Polymerase. Genetic screens in Neurospora , C. elegans , and Arabidopsis have identified several genes that appear to be crucial for on Biore, PTGS and RNAi. Several of these, including Neurospora qde-1 , Arabidopsis SDE-1/SGS-2 and C. elegans ego-1 , appear to encode RNA-dependent RNA polymerases (RdRPs). At first glance, it might be assumed that this is proof that an RdRP activity is required for RNAi. Certainly the existence of an boy overboard RdRP might explain the Pore Pack, remarkable efficiency of dsRNA-induced silencing if it amplifed either the dsRNA prior to cleavage or the siRNAs directly.
But mutants of accident rate these genes have varying phenotypes, which makes the role of Essay on Biore RdRP in RNAi difficult to discern (1, 3, 17, 18). In C. elegans ego-1 mutants (ego stands for enhancer of glp-1 ), RNAi functions normally in somatic cells, but is defective in accident rate in malaysia germline cells where ego-1 is primarily expressed. Essay On Biore Pore! In Arabidopsis SDE-1 / SGS-2 mutants (SGS stands for suppressor of gene silencing), siRNAs are produced when dsRNA is introduced via an endogenously replicating RNA virus, but not when introduced by a transgene. Accident In Malaysia! It has been proposed that perhaps the Pore, viral RdRP is substituting for the Arabidopsis enzyme in these mutants (1, 3, 17, 18). Although no homolog of an breakfast essay RdRP has been found in flies or humans, an RdRP activity has recently been reported in Drosophila embryo lysates (30). One model of amplification, termed the Essay on Biore, random degradative PCR model, suggests that an RdRP uses the guide strand of an siRNA as a primer for what, the target mRNA, generating a dsRNA substrate for Dicer and thus more siRNAs (27, 30). Evidence supporting this model has been found in on Biore worms, whereas experimental results refuting the model have been obtained from Drosophila embryo lysates (26, 27).
Two C. elegans genes, rde-1 and rde-4 (rde stands for RNAi deficient), are believed to be involved in the initiation step of RNAi. Mutants of these genes produce animals that are resistant to is quackery silencing by injection of dsRNA, but silencing can be effected in these animals by on Biore, the transmission of siRNA from heterozygous parents that are not silencing deficient. The C. elegans rde-1 gene is a member of a large family of genes and is homologous to the Neurospora qde-2 (qde stands for School Policy, quelling deficient) and on Biore Pack the Arabidopsis AGO1 genes (AGO stands for argonaute; AGO1 was previously identified to be involved in Arabidopsis development). Although the function of these genes in PTGS is unclear, a mammalian member of the RDE-1 family has been identified as a translation initiation factor. Interestingly, Arabidopsis mutants of AGO1 , which are defective for verses at Home Wives Essay, cosuppression, also exhibit defects in Essay leaf development.
Thus some processes or enzymes involved in PTGS may also be involved in development (1, 3, 17, 18). Important genes for the effector step of PTGS include the C. elegans rde-2 and mut-7 genes. These genes were initially identified from heterozygous mutant worms that were unable to transmit RNAi to their homozygous offspring (16). Worms with mutated rde-2 or mut-7 genes exhibit defective RNAi, but interestingly, they also demonstrate increased levels of accident transposon activity. Thus, silencing of transposons appears to occur by a mechanism related to RNAi and PTGS. Although the rde-2 gene product has not yet been identified, the mut-7 gene encodes a protein with homology to the nuclease domains of Essay on Biore RNase D and a protein implicated in Werner syndrome (a rapid aging disease) in humans (1, 3, 17, 18, 31). What! Perhaps this protein is a candidate for the nuclease activity required for target RNA degradation. PTGS Has Ancient Roots.
Discoveries from both genetic and biochemical approaches point to the fact that PTGS has deep evolutionary roots. Proposals have been put forth that PTGS evolved as a defense mechanism against transposons or RNA viruses, perhaps before plants and animals diverged (1, 3, 17, 18). Interestingly, it was noted by many researchers that disruption of genes required for RNAi often causes severe developmental defects. This observation suggested a link between RNAi and at Essay least one developmental pathway. A group of small RNA molecules, known as small temporal RNAs (stRNAs), regulates C. elegans developmental timing through translational repression of target transcripts. Research indicates that the C. Verses At Home! elegans lin-4 and let-7 stRNAs are generated from Essay on Biore Pore Pack 70-nt transcripts following the folding of these longer transcripts into in malaysia, a stem-loop structure. The folded RNA molecules are cleaved to produce 22-nt stRNAs by the enzyme Dicer (called DCR-1 in Essay Pack C. elegans ). Thus Dicer generates both siRNAs and stRNAs, and represents an regeneration (novel) intersection point for the RNAi and stRNA pathways (32-34). Recently, nearly 100 additional. 22 nt RNA molecules, termed microRNAs (miRNAs), were identified in Essay on Biore Pore Pack Drosophila , C. elegans , and Policy on Make-up HeLa cells (35-38). Essay Pore Pack! Much like lin-4 and let-7 , these miRNAs are formed from precursor RNA molecules that fold into a stem-loop secondary structure.
The newly discovered. 22 nt miRNAs are believed to play a role in regulation of gene expression, and at least two of them are known to require Dicer for their production (37). It appears that the use of small RNAs for both gene regulation and breakfast club essay RNAi is a common theme throughout evolution. Inducing RNAi in Mammalian Cells #151; From Mechanism to Application. Non-specific Gene Silencing by Long dsRNAs. While the natural presence of Essay on Biore Pore RNAi had been observed in a variety of organisms (plants, protozoa, insects, and what nematodes), evidence for Essay on Biore Pore, the existence of RNAi in mammalian cells took longer to what is quackery establish.
Transfection of long dsRNA molecules (30 nt) into most mammalian cells causes nonspecific suppression of gene expression, as opposed to on Biore the gene-specific suppression seen in School Board Policy on Make-up Essay other organisms. This suppression has been attributed to Essay an antiviral response, which takes place through one of two pathways. In one pathway, long dsRNAs activate a protein kinase, PKR. Activated PKR, in turn phoshorylates and inactivates the translation initiation factor, eIF2a, leading to repression of translation. (39) In the boy overboard, other pathway, long dsRNAs activate RNase L, which leads to nonspecific RNA degradation (40). A number of groups have shown that the dsRNA-induced antiviral response is absent from mouse embryonic stem (ES) cells and at least one cell line of embryonic origin. (41, 42) It is therefore possible to use long dsRNAs to silence specific genes in these specific mammalian cells.
However, the antiviral response precludes the use of long dsRNAs to induce RNAi in most other mammalian cell types. siRNAs Bypass the Antiviral Response. Interestingly, dsRNAs less than 30 nt in Essay on Biore Pore Pack length do not activate the PKR kinase pathway. This observation, as well as knowledge that long dsRNAs are cleaved to form siRNAs in worms and flies and that siRNAs can induce RNAi in Drosophila embryo lysates, prompted researchers to test whether introduction of breakfast club essay siRNAs could induce gene-specific silencing in mammalian cells (43). Indeed, siRNAs introduced by transient transfection were found to effectively induce RNAi in mammalian cultured cells in a sequence-specific manner. Pack! The effectiveness of siRNAs varies #151; the most potent siRNAs result in 90% reduction in target RNA and regeneration (novel) protein levels (44-46). The most effective siRNAs turn out to be 21 nt dsRNAs with 2 nt 3' overhangs. Sequence specificity of Essay Pore siRNA is very stringent, as single base pair mismatches between the siRNA and its target mRNA dramatically reduce silencing (44, 47). Unfortunately, not all siRNAs with these characteristics are effective. The reasons for this are unclear but may be a result of positional effects (46, 48, 49). For current recommendations on designing siRNAs, see siRNA Design.
RNAi as a Tool for accident rate, Functional Genomics. Although the Essay, history and mechanism of RNAi and PTGS are fascinating, many researchers are most excited about Policy, RNAi's potential use as a functional genomics tool. On Biore Pack! Already RNAi has been used to ascertain the function of many genes in Drosophila , C. elegans , and several species of plants. Boy Overboard! With the knowledge that RNAi can be induced in mammalian cells by the transfection of siRNAs, many more researchers are beginning to Pore Pack use RNAi as a tool in human, mouse and other mammalian cell culture systems. In early experiments with mammalian cells, the siRNAs were synthesized chemically (Ambion is one of several companies that offer custom siRNA synthesis). Stay Wives Essay! Recently, Ambion introduced a kit (the Silencer #153; siRNA Construction Kit) to produce siRNAs by in vitro transcription, which is Essay on Biore Pore Pack a less expensive alternative to Policy Work Essay chemical synthesis, particularly when multiple different siRNAs need to Essay Pore be synthesized. Verses At Home Wives! Once made, the siRNAs are introduced into cells via transient transfection. Due to differences in efficacy, most researchers will synthesize 3#150;4 siRNAs to a target gene and perform pilot experiments to determine the most effective one. Transient silencing of more than 90% has been observed with this type of approach (44-46, 48, 49).
So far, injection and transfection of dsRNA into cells and organisms have been the main method of delivery of siRNA. And while the silencing effect lasts for several days and does appear to be transferred to daughter cells, it does eventually diminish. Recently, however, a number of Essay groups have developed expression vectors to continually express siRNAs in transiently and stably transfected mammalian cells (50-56). Accident Rate! Some of these vectors have been engineered to express small hairpin RNAs (shRNAs), which get processed in vivo into siRNAs-like molecules capable of Essay on Biore Pore Pack carrying out gene-specific silencing (50, 53, 54, 56). The vectors contain the boy overboard, shRNA sequence between a polymerase III (pol III) promoter and a 4-5 thymidine transcription termination site. The transcript is terminated at position 2 of the Essay on Biore Pore Pack, termination site (pol III transcripts naturally lack poly(A) tails) and then folds into a stem-loop structure with 3' UU-overhangs. The ends of the shRNAs are processed in Wives verses Stay at Home Wives Essay vivo, converting the shRNAs into. 21 nt siRNA-like molecules, which in turn initiate RNAi (50). This latter finding correlates with recent experiments in C. elegans , Drosophila , plants and Trypanosomes, where RNAi has been induced by Essay Pore, an RNA molecule that folds into boy overboard, a stem-loop structure (reviewed in 3). Another siRNA expression vector developed by a different research group encodes the Pack, sense and regeneration (novel) antisense siRNA strands under control of separate pol III promoters (52). The siRNA strands from Pore Pack this vector, like the shRNAs of the other vectors, have 5 thymidine termination signals.
Silencing efficacy by boy overboard, both types of expression vectors was comparable to that induced by transiently transfecting siRNA. The recent studies on RNAi have taken the research world by storm. The ability to quickly and easily create loss-of-function phenotypes has researchers rushing to learn as much as they can about Essay on Biore Pore Pack, RNAi and the characteristics of effective siRNAs. In the future, RNAi may even hold promise for development of gene-specific therapeutics. Much has been learned about Board Policy Work, this powerful technique, but additional information becomes available on an almost daily basis (see The RNA Interference Resource to learn about the very latest RNAi research and tools). It is on Biore Pore Pack not an understatement to say that the accident in malaysia, field of functional genomics is being revolutionized by RNAi.
Cogoni C, and Macino G. (2000) Post-transcriptional gene silencing across kingdoms. Genes Dev 10 : 638-643. Essay Pore! (Abstract) Flash Animation: How Does RNAi Work? Hammond, S.M., Caudy, A.A., Hannon, G.J. Accident Rate In Malaysia! (2001) Post-transcriptional Gene Silencing by Double-stranded RNA. Nature Rev Gen 2 : 110-119. Warning : Failed opening '../../includes_new/ConnectDatabase.inc' for inclusion (include_path='.:/usr/local/lib/php') in /home/httpd/html/ambion/techlib/hottopics/rnai/rnai_may2002_print.html on Essay on Biore, line 723.
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